[1] PROTAS M E, PATEL N H.  Evolution of coloration patterns[J]. Annu Rev Cell Dev Biol, 2008, 24(1): 425-446.   doi: 10.1146/annurev.cellbio.24.110707.175302
[2] RODGERS G M, KELLEY J L, MORRELL L J.  Colour change and assortment in the western rainbowfish[J]. Anim Behav, 2010, 79(5): 1025-1030.   doi: 10.1016/j.anbehav.2010.01.017
[3] BRAASCH I, SCHARTL M, VOLFF J N.  Evolution of pigment synthesis pathways by gene and genome duplication in fish[J]. BMC Evol Biol, 2007, 7(1): 74-.   doi: 10.1186/1471-2148-7-74
[4] JIANG Y, ZHANG S, XU J, et al.  Comparative transcriptome analysis reveals the genetic basis of skin color variation in common carp[J]. PLoS One, 2014, 9(9): e108200-.   doi: 10.1371/journal.pone.0108200
[5] RAWLS J F, MELLGREN E M, JOHNSON S L.  How the zebrafish gets its stripes[J]. Dev Biol, 2001, 240(2): 301-314.   doi: 10.1006/dbio.2001.0418
[6] LIN J Y, FISHER D E.  Melanocyte biology and skin pigmentation[J]. Nature, 2007, 445(7130): 843-850.   doi: 10.1038/nature05660
[7] ALTSCHMIED J, DELFGAAUW J, WILDE B, et al.  Subfunctionalization of duplicate mitf genes associated with differential degeneration of alternative exons in fish[J]. Genetics, 2002, 161(1): 259-267.
[8] SUZUKI I, TADA A, OLLMANN M M, et al.  Agouti signaling protein inhibits melanogenesis and the response of human melanocytes to α-Melanotropin[J]. J Invest Dermatol, 1997, 108(6): 838-842.   doi: 10.1111/1523-1747.ep12292572
[9] TACHIBANA M.  MITF: a stream flowing for pigment cells[J]. Pigment Cell Res, 2000, 13(4): 230-240.   doi: 10.1034/j.1600-0749.2000.130404.x
[10]

章之蓉. 锦鲤[M]. 北京: 中国农业出版社, 2002: 2-31.

[11]

邹旭龙. 几种环境因子对锦鲤生长及体色影响的研究[D]. 大连: 大连海洋大学, 2016: 12-26.

[12] 张晨, 刘倩, 顾宪明, 等.  饲料中不同添加剂组合对锦鲤生长和体色的影响[J]. 大连海洋大学学报, 2017, 32(4): 410-415.
[13] 崔培, 姜志强, 王雪, 等.  不同蛋白水平的虾青素饲料对锦鲤体色、生长及免疫的影响[J]. 上海海洋大学学报, 2012, 21(3): 382-388.
[14] DAVID L, ROTHBARD S, RUBINSTEIN I, et al.  Aspects of red and black color inheritance in the Japanese ornamental (Koi) carp (Cyprinus carpio L.)[J]. Aquaculture, 2004, 233(1/2/3/4): 129-147.
[15] 刘启智, 肖军, 罗凯坤, 等.  雌核发育橙黄色锦鲤的遗传、性腺发育及外形特征[J]. 水产学报, 2013, 37(3): 390-396.
[16] LIU J H, WEN S, LUO C, et al.  Involvement of the mitfa gene in the development of pigment cell in Japanese ornamental (Koi) carp (Cyprinus carpio L.)[J]. Genet Mol Res, 2015, 14(1): 2775-2784.   doi: 10.4238/2015.March.31.7
[17] LI X, SONG Y, XIAO G, et al.  Gene expression variations of red-white skin coloration in common carp (Cyprinus carpio)[J]. Int J Mol Sci, 2015, 16(9): 21310-21329.   doi: 10.3390/ijms160921310
[18] LUO M K, WANG L M, ZHU W B.  Identification and characterization of skin color micro RNAs in Koi carp (Cyprinus carpio L.) by Illumina sequencing[J]. BMC Genomics, 2018, 19(1): 779-.   doi: 10.1186/s12864-018-5189-5
[19] 林光华, 翁世聪.  兴国红鲤的胚胎发育[J]. 南昌大学学报(理科版), 1986, (3): 1-8, 103.
[20] 许静, 谢从新, 邵俭, 等.  雅鲁藏布江尖裸鲤胚胎和仔稚鱼发育研究[J]. 水生态学杂志, 2011, 32(2): 86-95.
[21] 韩耀全, 何安尤, 蓝家湖, 等.  乌原鲤的胚胎发育特征[J]. 水产科学, 2018, 37(3): 368-373.
[22] 张哲, 卢毅, 詹俊阁, 等.  红白锦鲤的人工催产及胚胎发育研究[J]. 经济动物学报, 2017, 22(1): 1-6.
[23] 田雪, 庞小磊, 王良炎, 等.  MITFa及TYR基因在红色锦鲤体色发生不同阶段的表达分析[J]. 水产科学, 2017, 36(2): 197-201.
[24]

文胜. mitfa基因在观赏鱼体色形成中的功能研究[D]. 长沙: 湖南师范大学, 2014: 35-36.

[25] LECLERCQ E, TAYLOR J F, MIGAUD H, et al.  Morphological skin colour changes in teleosts[J]. Fish Fish, 2009, 11(2): 159-193.   doi: 10.1111/faf.2010.11.issue-2
[26] GOMELSKY B, CHERFAS N, HULATA G.  Studies on the inheritance of black patches in ornamental (Koi) carp[J]. Isr J Aquacult-Bamid, 1998, 50(3): 134-139.
[27] UYEN L D, NGUYEN D H, KIM E K.  Mechanism of skin pigmentation[J]. Biotechnol Bioproc E, 2008, 13(4): 383-395.   doi: 10.1007/s12257-008-0143-z
[28] HOEKSTRA H E.  Genetics, development and evolution of adaptive pigmentation in vertebrates[J]. Heredity, 2006, 97(3): 222-234.   doi: 10.1038/sj.hdy.6800861
[29] ITO S, WAKAMATSU K.  Human hair melanins: what we have learned and have not learned from mouse coat color pigmentation[J]. Pigm Cell Melanoma R, 2011, 24(1): 63-74.   doi: 10.1111/j.1755-148X.2010.00755.x
[30] ZOU J, BEERMANN F, WANG J X, et al.  The fugu tyrp1 promoter directs specific GFP expression in zebrafish: tools to study the RPE and the neural crest-derived melanophores[J]. Pigm Cell Melanoma R, 2006, 19(6): 615-627.   doi: 10.1111/pcr.2006.19.issue-6
[31] VOISEY J, CARROLL L, van DAAL A.  Melanocortins and their receptors and antagonists[J]. Curr Drug Targets, 2003, 4(7): 586-597.   doi: 10.2174/1389450033490858
[32] PARICHY D M.  Evolution of danio pigment pattern development[J]. Heredity, 2006, 97(3): 200-210.   doi: 10.1038/sj.hdy.6800867
[33] CARREIRA S, GOODALL J, DENAT L, et al.  Mitf regulation of Dia1 controls melanoma proliferation and invasiveness[J]. Genes Dev, 2006, 20(24): 3426-3439.   doi: 10.1101/gad.406406
[34] THOMAS A J, ERICKSON C A.  The making of a melanocyte: the specification of melanoblasts from the neural crest[J]. Pig Cell Melanoma R, 2008, 21(6): 598-610.   doi: 10.1111/pcr.2008.21.issue-6
[35] ZHU W B, WANG L M, DONG Z J, et al.  Comparative transcriptome analysis identifies candidate genes related to skin color differentiation in red tilapia[J]. Sci Rep, 2016, 6(1): 31347-.   doi: 10.1038/srep31347
[36] 王巍, 胡红霞, 孙向军, 等.  锦鲤酪氨酸酶基因序列分析及其在不同锦鲤品系不同组织中的表达[J]. 水产学报, 2012, 36(11): 1658-1666.